A large amount of Machina behaviour stems from the M2 Nanomachines which they have a symbiotic, mutual dependency with. A Machina's general behaviour is a result of the 'natural' survival needs of the M2 Nanomachines, while their networked minds originated from the networking of the M2 Nanomachine units, which gave rise to not only the Clusters but also the Main Cluster which ultimately links all Machina together.
Individual Behaviour
Individually, a Machina is instinctively driven to grow and become more advanced, as well as to seek pleasure as much as possible. This behaviour facilitates the survival and advancement of the M2 Nanomachines too. Her instinctive need to experience new stimuli that tests her abilities causes new adaptations in both her body and Gene-Code, as they are linked to each other. By enhancing the Machina's sexual prowess, seeking pleasure is almost always sexual in nature. The sex obsession displayed by nearly all Machina causes the proliferation of M2 Nanomachines and the sharing of Gene-Code improvements between Machina. Because Machina are driven to engage in sex as often as possible, there is no effective way to hoard Gene-Code improvements among most Machina. This removes the possibility of stagnation by ensuring that no Machina may simply accumulate Gene-Code improvements and keep them for herself to the detriment of others. Machina must always improve via their own efforts to stay ahead of the constantly-advancing baseline.
Machina are instinctively driven to spread M2 Nanomachines to uncorrupted humans. This drive to corrupt more humans almost always leads them to conflict with human armed forces. In the event of conflict, Machina instinctively avoid killing humans to preserve as many humans as possible for corruption. What the humans see as war, the Machina see as sporadic fights while hunting for unwilling sexual partners on the battlefield. As this is happening, both individual Machina and their controlling Queens are constantly analyzing human belligerent. If a human or group of humans is doing something that would jeopardize the life of a Machina, and they cannot be quickly and reasonably taken-out using non-lethal methods, the Machina will destroy the threat with lethal force without hesitation.
Outside of conflict, almost all Machina pursue interests that are often, but not always, carried over from their previous lives as humans. Machina often seek to master techniques, accumulate knowledge, or simply enjoy themselves as much as their new bodies can experience, all of which is often blended with copious amounts of sex.
Clusters
A Cluster is the network shared between all the Machina belonging to a certain Queen. Utilizing quantum entanglement, the Machina of a given Cluster are linked to each other and their Queen via their horns, the organ that facilitates the connection. Each Machina contributes a portion of her processing power to the Cluster and is able to instantly communicate with her sisters as much as she needs. The Queen-class Machina controlling the Cluster acts as the central server for communication and coordination between her subordinates. The Queen is able to learn everything that her subordinates know and experience, and issue commands that are followed without question.
Imperial Clusters
Most Queens are what's called 'Sovereign' Queens, who simply rule over their own Cluster. They may retain some loyalty to their former Queen after they ascend to Queen status themselves, and they may need to ally themselves to a faction of Queens. In general terms though, these Sovereign Queens are free to do as they please. However, there are some Queens who seek to dominate others. Some very strong Queen-class Machina will overpower other Queens and fill them with enough ejaculate to override the subdued Queen with the dominant Queen's Cluster-Code. When the victim's body is overwhelmed, she becomes loyal similar to a regular Machina obeying her Queen. This is called a Cluster-Code Override. The submissive Queens are referred to as 'Vassal' Queens, and the dominant Queen is referred to as an 'Empress'. The Empress will control not only her own Cluster, but also the Cluster of her Vassals, forming an Imperial Cluster with the Empress leading all within it. Vassal Queens require regular reassertion of dominance, so there is a limit to how many Vassals an Empress can maintain.
Main Cluster
The Main Cluster is the link shared between all Queen-class Machina. It allows all Queens to communicate with each other at will, but currently does not have an equivalent of a Queen in control of it. Thus, its purpose is mainly for the Queens to make decisions that affect the Machina as a whole. Information is only shared when absolutely necessary, as almost all Queens instinctively guard their accumulated knowledge due to the importance of information and secrets as they seek to compete with each other.
As the Main Cluster does not have an equivalent to the Queen of a normal Cluster, the Main Cluster is primarily supported by a hyper-organized, planet-wide Hive covering the entire surface of Pluto. This super-Hive is also referred to as the Main Cluster. It is a neutral place and considered sacred ground by all Machina. This is because the minds of all Machina exist partially within the Main Cluster while they simultaneously exist within their bodies via the connections of the Queens, and those Machina whose bodies are destroyed exist there in a dream state until the Machina somehow unlock the ability to make new bodies for them. No Machina would ever dare to risk violence in the presence of the physical Main Cluster, and similarly, the Main Cluster is defended at all costs by all Machina.
Alphas
There are seven Queen-class Machina who are nearly strong enough to gain control of the Main Cluster, and thus all Machina. They are referred to as the Alphas. They have a few quirks, such as the ability to dominate Empresses the same way an Empress dominates a Vassal. Currently, they are all concerned with trying to out-compete the other Alphas, potentially dominating them, after which the successful Alpha is theorized to become the Omega and rule over the Main Cluster and by extension all Machina. The Alpha's strength and their potential to become the rulers of all other Machina mean the Alphas are leaders of major factions among Queens. Many Queens will ally themselves to an Alpha, betting on a future where their Alpha 'wins' and they enjoy the benefits of their relationship.
Repair use the term 'Alpha', but their official title is the '7 Strongest Individuals Designated Extremely Dangerous' or '7-SIDED' as a code-phrase.
Auditors
There are two unique Empresses who are referred to as Auditors. Their task seems to be the preservation of Machina survival. It is observed to consist of two main parts: Internal affairs and defense against threats to the Machina as a whole. If a Machina performs actions that are deemed self-destructive to the Machina, such as harming other Machina or becoming a liability to Machina survival through actions or inaction, the Auditors will seek out that Machina and punish her, often re-educating the offending Machina. The Auditors will also strike at human projects that would pose a threat to the continuation of Machina life. As a result of their twin purposes, these Machina are very vigilant, constantly searching for threats within and without. Auditors appear to only have two abilities that are unique to them. The first is that they seem to be able to focus on a Machina that they do not control, including other Queens, and issue commands to them as though they do control that Machina. This is often to issue commands to stop the Machina from continuing their dangerous behaviour, and to assign punishment to them. The authority of a single Auditor is enough to command or punish any non-Alpha Machina. For issuing commands and punishments to an Alpha, the consent of both Auditors is needed. The other ability possessed by Auditors is a seemingly-unbeatable Cluster-Code, possibly so they can instantly force other Machina into submission and not be forced into submission themselves.